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Sunday, September 5, 2021

09-05-2021-0015 - Stramenopile

Stramenopile is a clade of organisms distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, and in some they have been secondarily lost (in which case relatedness to stramenopile ancestors is evident from other shared cytological features or from genetic similarity). Stramenopiles represent one of the three major clades in the SAR supergroup, along with Alveolata and Rhizaria.

Members of the clade are referred to as 'stramenopiles'. Stramenopiles are eukaryotes; since they are neither fungi, animals, nor plants, they are classified as protists. Most stramenopiles are single-celled, but some are multicellular algae including some brown algae. The group includes a variety of algal protists, heterotrophic flagellates, opalines and closely related proteromonad flagellates (all endobionts in other organisms); the actinophryid heliozoa, and oomycetes. The tripartite hairs have been lost in some stramenopiles - for example in most diatoms (although these organisms still express mastigonemic proteins - see below).

Many stramenopiles are unicellular flagellates, and most others produce flagellated cells at some point in their lifecycles, for instance as gametes or zoospores. Most flagellated heterokonts have two flagella; the anterior flagellum has one or two rows of stiff hairs or mastigonemes, and the posterior flagellum is without such embellishments, being smooth, usually shorter, or in a few cases not projecting from the cell. 

Electron micrograph of the protist Paraphysomonas butcheri. It illustrates the stramenopile property - of having stiff hairs. The hairs attach to one longer flagellum, the other is without hairs (an arrangement also called 'heterokont'). The body of the flagellate is coated with delicate scales. Paraphysomonas feeds on bacteria, two of which lie near the hairy flagellum.

https://en.wikipedia.org/wiki/Stramenopile

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